Typically paid (and fired) on marked to market.
Having MUCH experience with Chicago and to some extent NYC proprietary futures trading firms, I can say in terms of my own personal observations that I have never seen a trader get paid out anything close to his or her full share on MtoM, but I sure as hell have seen traders get instantly fired and escorted out of a building on MtoM. (I was enlisted by management to sort through and liquidate a few of these burning shit piles on occasion). It's part of the business.
I have seen some traders, myself included, get very healthy draws based upon protracted performance and positive equity. Every firm that I was ever at had pretty tight risk controls and sharp Risk Managers. Most did not like options trading unless there was a large element of market making or delta neutral spread arbitrage involved. Again, my own observations - YMMV.