Quote from john dough:
Changes in alleles do not necessarily carry with them a certificate of infertility. So, if a change in an allele in group AA is propagated to group AA offspring and does not affect intra-group fertility, then the change is preserved.
Similarly, for group BB. Only the change to group BB is almost certainly likely to be to a different allele.
At this point we have two groups, AA and BB, with two different genetic differences, likely appearing in two different loci in their respective genomes.
Now repeat this process, and keep repeating it until eventually, you will reach a point where the two genomes of AA and BB are sufficiently different so that the group members can no longer successfully mate inter-group.
Thus, they are speciated. Which is what the cited article demonstrates.
There is no such process without species A becoming mutually infertile with at least one of its descendant species AA or BB.
So again, you're dodging the question.